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Adaptations for water. Anchored in one place, plants
can't travel to search for life-sustaining water. Plants growing in regions
where water is scarce or available only during a brief rainy season have
adopted various strategies for survival. Let's voyage into a desert, and
look at the various strategies for survival in an environment where water
is in short supply.
Think of a desert, and most likely what first comes to mind is the cactus.
Cacti are among the most drought-resistant plants on earth. What strategies
have cacti evolved to endure desert conditions? Here are a few characteristics
common to many cactus species:
- Succulent stems allow cacti to store water. The barrel cactus has
an accordion-like stem that allows the cactus to swell as the roots
take in water during a rainy spell, then shrink as the moisture is used
up.
- The spines on a cactus perform a number of important functions. They
help to shade the plant from intense sun, helping to cool the surface.
And, if you've ever bumped into a cactus by mistake, you know that they
also serve as protection.
- An extensive, shallow root system allows cacti to take advantage of
even the lightest rainfall.
- During the hot, dry summer, many desert plants drop their leaves and
become dormant. Cactus spines are modified leaves; photosynthesis takes
place in the stem and can continue throughout the summer. With no leaves,
the surface area exposed to the sun and drying winds is reduced, which
in turn reduces water loss from transpiration.
- Cacti have a waxy cuticle layer that helps seal in moisture.
Here are some other ways desert plants cope with a scarcity of water:
- The desert lily's bulbs are found at a depth of 12 to 18 inches. The
plant can lie dormant for years until adequate water is available.
- Welwitchia, a desert gymnosperm, absorbs water from the nighttime
fog through its leaf stomata.
- The creosote bush has a far-ranging root system: roots may reach 75
feet in length.

- Some plants complete their life cycles in months, or even weeks. Desert
sand verbena seeds germinate in the spring after the winter rains. They
grow quickly, bloom, and produce and scatter seeds, then die with the
onset of summer heat. The seeds are extremely durable and drought-resistant,
and may lie dormant for years until conditions are favorable for germination.
Plants that bloom for a brief time in the spring are often referred
to as ephemerals.
Scarcity of water isn't restricted to desert environments. Even some rainforest
plants are challenged to meet their water needs. We mentioned in the section
above that certain rainforest plants, called epiphytes, grow in the branches
of the canopy. Lacking the ability to draw water from the soil, they have
evolved other strategies: 
Orchids have aerial roots that can absorb atmospheric water vapor.

Bromeliads grow in tree branches; their leaves grow in cup-shaped rosettes
that collect rainwater.
Closer to the home front, you have probably seen a dull, powdery, bluish-white
coating on various fruits, including plums, apples, and grapes. This natural
surface wax, called a "bloom," is easily wiped off.
Other plants with a noticeable bloom include some succulents and cacti;
also the leaves of carnations and cabbages (especially red cabbage). Although
all plants have a cuticle layer, this visible bloom occurs when
the surface wax occurs in tiny "rodlets" that protrude from the surface
cells. Like all cuticles, the bloom helps conserve water by reducing evaporation
from surface tissues. The word glaucous
is used to describe plants with a visible waxy bloom.
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